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Willson MF, Womble JN. 2006. Vertebrate exploitation of pulsed marine prey: a review and the example of spawning herring. Reviews in Fish Biology and Fisheries. 16:183-200.

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Womble JN, Sigler MF, Willson MF. 2009. Linking seasonal distribution patterns with prey availability in a central-place forager, the Steller sea lion. Journal of Biogeography. 36:439-451.

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Womble JN, Willson MF, Sigler MF, Kelly BP, VanBlaricom GR. 2005. Distribution of Steller sea lions Eumetopias jubatus in relation to spring-spawning fish in SE Alaska. Marine Ecology Progress Series. 294:271-282.

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Gende SM, Womble JN, Willson MF, Marston BH. 2001. Cooperative foraging by Steller sea lions. Canadian Field Naturalist. 115:355-356.